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Hemerobiidae sp.

Hemerobiidae sp.


Family: Hemerobiidae
Order: Neuroptera
Class: Insecta
Phylum / Division: Arthropoda
Kingdom: Animalia

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Description


Hemerobiidae is a family of Neuropteran insects commonly known as brown lacewings, comprising about 500 species in 28 genera. Most are yellow to dark brown, but some species are green. They are small; most have forewings 4–10 mm long (some up to 18 mm). These insects differ from the somewhat similar Chrysopidae (green lacewings) not only by the usual coloring but also by the wing venation: hemerobiids differ from chrysopids in having numerous long veins (two or more radial sectors) and forked costal cross veins. Some genera (Hemerobius, Micromus, Notiobiella, Sympherobius, Wesmaelius) are widespread, but most are restricted to a single biogeographical realm. Some species have reduced wings to the degree that they are flightless. Imagines (adults) of subfamily Drepanepteryginae mimic dead leaves. Hemerobiid larvae are usually less hairy than chrysopid larvae.

Hemerobiids, like chrysopids, are predatory, especially on aphids, both as larvae and adults. The species Micromus tasmaniae is bred for biological pest control.

Systematics: The relationships between the Hemerobiidae and the other Neuropteran families are still unresolved. Despite their superficial similarity to chrysopids, the brown and green lacewings may not be as closely related as was at one time believed. Rather, the Hemerobiidae appear to be closely related to the Mantispidae.

The superfamily Hemerobioidea is currently restricted to the Hemerobiidae. Formerly, the pleasing lacewings (Dilaridae), silky lacewings (Psychopsidae), giant lacewings (Polystoechotidae) and as noted above the green lacewings (Chrysopidae) were placed therein too. Of these, only the Dilaridae and Chrysopidae seem to be reasonably close relatives of the brown lacewings. The Psychopsidae in fact seem to belong to an altogether different suborder of Neuroptera, the Myrmeleontiformia.

Phylogeny: Cladogram of Hemerobiidae relations, based on morphological and molecular data. Psychobiellinae was rearranged into Notiobiellinae and Zachobiellinae, and Adelphohemerobiinae was placed as incertae sedis.

Genera: The subfamilies of Hemerobiidae are:

  • Subfamily Drepanacrinae
  • Genus Austromegalomus
  • Genus Conchopterella
  • Genus Drepanacra
  • Genus Psychobiella
  • Subfamily Carobiinae
  • Genus Carobius
  • Subfamily Drepanepteryginae
  • Genus Drepanepteryx Leach, 1815
  • Genus Gayomyia
  • Genus Neuronema
  • Subfamily Hemerobiinae
  • Genus Biramus
  • Genus Hemerobiella
  • Genus Hemerobius Linnaeus, 1758
  • Genus Nesobiella
  • Genus Wesmaelius Krüger, 1922
  • Subfamily Megalominae
  • Genus Megalomus Rambur, 1842
  • Subfamily Microminae
  • Genus Megalomina Banks, 1909
  • Genus Micromus Rambur, 1842
  • Genus Nusalala Navás, 1913
  • Subfamily Notiobiellinae
  • Genus Notiobiella
  • Subfamily Sympherobiinae
  • Genus Neosympherobius
  • Genus Nesobiella
  • Genus Sympherobius Banks, 1904
  • Subfamily Zachobiellinae
  • Genus Anapsectra
  • Genus Psectra Hagen, 1866
  • Genus Zachobiella
  • Subfamily incertae sedis
  • Genus Adelphohemerobius Oswald, 1993
  • Genus Notherobius - uncertain or fairly basal position


Fossils: Numerous fossil Hemerobiidae have been described, some from the still-living genera, others from genera that are entirely extinct today. While most have been found in Eocene to Miocene rocks or amber, Promegalomus is known from the Jurassic. It was formerly considered to constitute a distinct family Promegalomidae, but is currently recognized as a very basal member of the Hemerobiidae. The Cretaceous Mesohemerobius was formerly considered a brown lacewing, but is today rather placed as incertae sedis in the Neuroptera; it might be a member of the Hemerobioidea but not even that is certain. Notable fossil Hemerobiidae genera are:
  • Bothromicromus Scudder, 1878 (Eocene/Oligocene; Quesnel, British Columbia)
  • Brasilopsychopsis Crato Formation Brazil, Early Cretaceous (Aptian)
  • Cratopsychopsis Crato Formation Brazil, Aptian
  • Cretomerobius Ponomarenko, 1992 Dzun-Bain Formation, Mongolia, Aptian
  • Hemeroberotha Makarkin & Gröhn, 2020 Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
  • Mucropalpus Pictet, 1856 (Eocene; Baltic amber)
  • Plesiorobius Klimaszewski and Kevan 1986 Late Cretaceous (Taimyr amber, Russia, Santonian Canadian amber, Campanian Ola Formation, Russia, Campanian)
  • Prochlanius Kruger, 1923 (Eocene; Baltic amber)
  • Promegalomus Panfilov, 1980 Karabastau Formation, Kazakhstan, Middle/Late Jurassic (Callovian/Oxfordian)
  • Prophlebonema Kruger, 1923 (Eocene; Baltic amber)
  • Prospadobius Kruger, 1923 (Eocene; Baltic amber)
  • Purbemerobius Jepson et al. 2012 Durlston Formation, United Kingdom, Early Cretaceous (Berriasian)

The extinct genus Hemerobites was originally described in 1813 from a specimen preserved in Baltic amber as a hemerobiid. However further study has resulted in its synonymy with the genus Eutermes and placement in the termite subfamily Termitina. The species Wesmaelius mathewesi, was described in 2003 from a solitary Eocene fossil found near Quesnel, British Columbia, and placed into the extant genus Wesmaelius. At that time it was the oldest Hemerobiinae species described. Another species from the same genus, Wesmaelius makarkini was found in Garang Formation of Zeku County, Qinghai Province, China, in 2018. The finding is from Lower Miocene.

Some additional brown lacewing larvae have been found as fossils, but it has been impossible to determine their generic or subfamilial association.

Source: Wikipedia, article Hemerobiidae sp.. Content adapted from Wikipedia, available under the CC BY-SA 4.0 license. Accessed on 17/07/2026.





Geographic distribution


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Species photographs

Count: 8

Foto
Photography ID: 659499
  Adult

Conesa
Río Negro
Argentina
11/14/2025
Miguel Angel Barberis
Foto
Photography ID: 623722
  Adult

Piriápolis
Maldonado
Uruguay
02/28/2025
Enrico H. Vicentini
Foto
Photography ID: 564792
  Adult

Florida
Buenos Aires
Argentina
11/10/2018
Marcos Garberoglio
Foto
Photography ID: 428529
  Adult

Santo Tomé
Santa Fe
Argentina
01/04/2021
Eduardo Beltrocco
Foto
Photography ID: 420723
  Adult

Santo Tomé
Santa Fe
Argentina
05/16/2020
Eduardo Beltrocco
Foto
Photography ID: 418902
  Adult

Santo Tomé
Santa Fe
Argentina
05/19/2020
Eduardo Beltrocco
Foto
Photography ID: 415796
  Adult

Santo Tomé
Santa Fe
Argentina
09/25/2020
Eduardo Beltrocco


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Species records

Count: 8



Page 1 of 1
Record IDDateExact timeCountryProvince / departmentPlaceFilmedPhotographedRecorded vocalObservedHeardWounded or deadNumber of individualsUser or BibliographyDetail
240315614/11/2025ArgentinaRío NegroGeneral Conesa, ConesaMiguel Angel Barberis
225491028/02/2025UruguayMaldonadoPunta Negra, PiriápolisEnrico H. Vicentini
219023414/11/2024ArgentinaCiudad Autónoma de Buenos AiresReserva Ecológica Ciudad Universitaria - Costanera Norte (RECU-CN)Marcos Garberoglio
134736504/01/2021ArgentinaSanta FeAvenida Luján, entre Gaboto y Colón, Santo ToméEduardo Beltrocco
130261825/09/2020ArgentinaSanta FeSarmiento y Centenario, Santo ToméEduardo Beltrocco
124670919/05/2020ArgentinaSanta FeSarmiento y Centenario, Santo Tomé1Eduardo Beltrocco
131608216/05/2020ArgentinaSanta FeSarmiento y Centenario, Santo ToméEduardo Beltrocco
194827610/11/2018ArgentinaBuenos AiresFloridaMarcos Garberoglio
Page 1 of 1

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Recommended citation
Please use the following reference when citing this species account.
EcoRegistros. 2026. Hemerobiidae sp. – Species account. Accessed from www.ecoregistros.org on 14/09/2026.